Dru tells Gru about their father, one of the greatest villains of all-time. They talk about their family tradition of being a villain, and Dru asks Gru to teach him to be a villain since he is considered a failure. Gru declines the offer, as he has left that life behind, but he takes a ride with Dru in the villain vehicle after Dru's tempting presentation of it.
Olympia and Otis attempt to intercept a dancing villain, who makes her victims dance forever. The villain strikes at different parts of town, and the duo guessed her attack pattern incorrectly. They determine that she will strike the mayor's home next after discovering the real pattern.
The CyberSquad is having trouble keeping track of Buzz and Delete's moves, so Inez suggests they determine the floor that Buzz and Delete are on and go directly there instead of following each individual move. She draws a number line to make it easier to find out where the two villains are. The villains' most recent moves confuse the group, as they end up below the ground floor. Matt, Inez, and Jackie realize that they need to travel to negative floor numbers and define negative numbers for the confused Digit.